1. Department of Earth and Planetary Science, the University of Tokyo, 7-3-1, Hongo, Bunkyo, Tokyo 113-0033, Japan
2. Kanagawa Prefectural Museum of Natural History, 499, Odawara, Kanagawa 250-0031, Japan (e-mail: )
3. Faculty of Environment and Information Sciences, Yokohama National University, 79-7, Tokiwadai, Hodogaya, Yokohama, Kanagawa 240-8501, Japan
4. Department of Cell and Developmental Biology, Biosciences, University College London, Gower Street, London, WC1E 6BT, UK