1. Zoological Laboratory, Okinawa Churashima Research Center, Okinawa Churashima Foundation, 888 Ishikawa, Motobu-cho, Okinawa 905-0206, Japan (e-mail: )
2. Kitakyushu Museum of Natural History and Human History, 2-4-1 Higashida, Yahatahigashi-ku, Kitakyushu, Fukuoka 805-0071, Japan
3. Sogokaihatsu Co., Ltd., 440-11, Oshiba, Mobara, Chiba 297-0033, Japan