1. a Mammal Research Institute , University of Pretoria , Pretoria , South Africa
2. b Department of Environmental Studies , Sweet Briar College , Sweet Briar , USA
3. c Mammal Research Institute , University of Pretoria , Pretoria , South Africa
4. d South African National Parts and Endangered Wildlife Trust , Skukuza , South Africa
5. e Samburu-Laikipia Wild Dog Project , Mpala Research Centre , Nanyuki , Kenya