1. Central Forest State Nature Biosphere Reserve, 172521, Zapovedniy Village, Russia
2. Joint Directorate of the Mordovia State Nature Reserve and National Park “Smolny”, 430005, Saransk, Russia
3. Scientific Research Zoological Museum, Moscow Lomonosov State University, 125009, Moscow, Russia
4. Biodiversity Research Institute (IMIB, CSIC/University of Oviedo/Principality of Asturias), Campus Mieres, 33600, Mieres, Spain
5. Department of Evolutionary Ecology, National Museum of Natural Sciences (MNCN), Spanish National Research Council (CSIC), 28006, Madrid, Spain