1. Department of Natural Resources, University of Mohaghegh Ardabili, Daneshgah Street, Ardabil, Iran
2. Department of Plant Sciences and Medicinal Plants, University of Mohaghegh Ardabili, Faculty of Agriculture, Meshgin-Shahr Campus, Daneshgah Street, Meshgin Shahr, Ardabil Province, Iran
3. Watershed Management, Forests and Rangelands Research Department, Guilan Agricultural and Natural Resources Research and Education Center, Agricultural Research, Education and Extension Organization (AREEO), Tehran Highway, Rasht, Guilan, Iran