1. Key Laboratory of Alpine Ecology and Biodiversity, Institute of Tibetan Plateau Research, Chinese Academy of Sciences, No.16 Lincui Road, Chaoyang District, Beijing 100101, China
2. Institute of Botany, Czech Academy of Sciences, Zamek 1, 252 43 Průhonice, Czech Republic
3. Department of Botany, Faculty of Science, Charles University, Benatska 2, 128 01 Prague, Czech Republic