1. State Key Laboratory of Tropical Oceanography, South China Sea Institute of Oceanology, Chinese Academy of Sciences, Guangzhou 510301, China
2. Daya Bay Marine Biology Research Station, Chinese Academy of Sciences, Shenzhen 518121, China
3. Innovation Academy of South China Sea Ecology and Environmental Engineering, Chinese Academy of Sciences, Guangzhou 510301, China
4. University of Chinese Academy of Sciences, Beijing 100049, China