1. State Key Laboratory of Biobased Material and Green Papermaking (LBMP), Qilu University of Technology (Shandong Academy of Sciences), Jinan 250353, China
2. Key Laboratory of Shandong Microbial Engineering, College of Bioengineering, Qilu University of Technology (Shandong Academy of Sciences), Jinan 250353, China
3. Gansu Engineering Technology Research Center for Microalgae, Hexi University, Zhangye 734000, China