1. Sea Search Research and Conservation, Muizenberg, Cape Town 7945, South Africa
2. School of Life Sciences, Department of Agriculture, Engineering and Science, University of KwaZulu-Natal, Durban 3629, South Africa
3. Department of Botany and Zoology, Faculty of Science, Stellenbosch University, Cape Town 7605, South Africa
4. Centre for Statistics in Ecology, Environment and Conservation, Department of Statistical Sciences, University of Cape Town, Cape Town 7701, South Africa