1. Graduate School of Horticulture, Chiba University, Matsudo 648, Matsudo-shi 271-8510, Japan
2. Graduate School of Life Sciences, Department of Ecological Developmental Adaptability Life Sciences, Tohoku University, 2-1-1 Katahira, Sendai 980-8577, Japan
3. Suntory Rising Stars Encouragement Program in Life Sciences (SunRiSE), Sendai 980-8578, Japan
4. Plant Molecular Science Center, Chiba University, 1-33 Yayoi, Chiba-shi 263-8522, Japan