1. Kaust Beacon Development, King Abdullah University of Science and Technology, Thuwal, Jeddah 23955, Saudi Arabia
2. Oceanography Department, Faculty of Science, Alexandria University, Alexandria 5424041, Egypt
3. Ministry of Environment, Water and Agriculture, King AbdulAziz Rd., Riyadh 11195, Saudi Arabia
4. Institute of Aquaculture, University of Stirling, Stirling FK9 4LA, UK