1. Independent Researcher, Woking GU21 5TR, UK
2. Department of Evolutionary Ecology, Leibniz Institute for Zoo and Wildlife Research, Alfred-Kowalke-Straße 17, 10315 Berlin, Germany
3. Wildlife Conservation Research Unit, Department of Biology, University of Oxford, The Recanati-Kaplan Centre, Tubney House, Abingdon Road, Abingdon OX13 5QL, UK
4. Department of Chemistry and Bioscience, Aalborg University, Fredrik Bajers Vej 7H, DK-9220 Aalborg, Denmark
5. Linacre College, St. Cross Road, Oxford OX1 3JA, UK