1. Key Laboratory of Plant Resource Conservation and Germplasm Innovation in Mountainous Region (Ministry of Education), College of Life Sciences/Institute of Agro-Bioengineering, Guizhou University, Guiyang 550025, China
2. Research Center for Biodiversiey and Nature Conservation, College of Forestry, Guizhou University, Guiyang 550025, China
3. Guizhou Chishui Alsophila National Nature Reserve Administration Bureau, Chishui 564700, China