1. Laboratory of Marine Biology and Ecology, Third Institute of Oceanography, Ministry of Natural Resources, Xiamen 361005, China
2. College of Marine Sciences, Shanghai Ocean University, Shanghai 201306, China
3. Scientific Observing and Experimental Station of Oceanic Fisheries Resources and Environment, Ministry of Agriculture, Shanghai 201306, China
4. School of Marine Sciences, University of Maine, Orono, ME 04469, USA