1. Institute of Zoology, Zoological Society of London, Regent’s Park, London NW1 4RY, UK
2. Independent Researcher, Purleigh CM3 6PZ, UK
3. Conservation and Policy, Zoological Society of London, Regent’s Park, London NW1 4RY, UK
4. Environment Agency, Rivers House, Threshelfords Business Park, Inworth Road, Feering CO5 9SE, UK