1. South China Sea Fisheries Research Institute, Chinese Academy of Fisheries Sciences, Guangzhou 510300, China
2. School of Fisheries, Zhejiang Ocean University, Zhoushan 316022, China
3. Key Laboratory for Sustainable Utilization of Open-Sea Fishery, Ministry of Agriculture and Rural Affairs, Guangzhou 510300, China
4. Research Center of Fisheries Resource and Environment, Chinese Academy of Fishery Sciences, Beijing 100141, China
5. Key Laboratory of Marine Ranching, Ministry of Agriculture and Rural Affairs, Guangzhou 510300, China