1. College of Marine Science and Biological Engineering, Qingdao University of Science and Technology, Qingdao 266042, China
2. Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Laboratory for Marine Fisheries Science and Food Production Processes, Pilot National Laboratory for Marine Science and Technology (Qingdao), Key Laboratory of Maricultural Organism Disease Control, Ministry of Agriculture and Rural Affairs, Qingdao Key Laboratory of Mariculture Epidemiology and Biosecurity, Qingdao 266071, China
3. Key Laboratory of Animal Epidemiology and Zoonosis, Ministry of Agriculture, College of Veterinary Medicine, China Agricultural University, Beijing 100193, China
4. College of Fisheries and Life Science, Shanghai Ocean University, Shanghai 201306, China
5. Shanghai Institute of Virology, Shanghai Jiao Tong University School of Medicine, Shanghai 200025, China