1. Key Laboratory of National Forestry and Grassland Administration on Subtropical Forest Biodiversity Conservation, Co-Innovation Center for Sustainable Forestry in Southern China, College of Biology and the Environment, Nanjing Forestry University, Nanjing 210037, China
2. State Key Laboratory of Biocontrol and Guangdong Provincial Key Laboratory of Plant Resources, School of Life Sciences, Sun Yat-Sen University, Guangzhou 510275, China
3. Administrative Commission of Danxiashan National Park, Shaoguan 512300, China