1. School of Biological Engineering, Dalian Polytechnic University, Dalian 116034, China
2. Shandong Provincial Key Laboratory of Synthetic Biology, Key Laboratory of Biofuels, Qingdao Institute of Bioenergy and Bioprocess Technology, Chinese Academy of Sciences, Qingdao 266101, China
3. Sinopec Dalian Research Institute of Petroleum and Petrochemicals, Dalian 116045, China