1. Henan Key Laboratory of Ion-Beam Bioengineering, School of Physics, Zhengzhou University, Zhengzhou 450052, China
2. Plateau Livestock Genetic Resources Protection and Innovative Utilization Key Laboratory of Qinghai Province, Key Laboratory of Animal Genetics and Breeding on Tibetan Plateau, Ministry of Agriculture and Rural Affairs, Qinghai Academy of Animal and Veterinary Medicine, Qinghai University, Xining 810016, China
3. School of Agricultural Sciences, Zhengzhou University, Zhengzhou 450052, China
4. Haidong City Animal Disease Prevention and Control Center, Haidong 810600, China
5. Japan International Research Center for Agricultural Sciences, Crop, Livestock and Environment Division, Tsukuba 305-8686, Japan