1. Key Laboratory of Coastal Biology and Bioresource Utilization, Yantai Institute of Coastal Zone Research, Chinese Academy of Sciences, Yantai 264003, China
2. University of Chinese Academy of Sciences, Beijing 100049, China
3. Center for Ocean Mega-Science, Chinese Academy of Sciences, Qingdao 266071, China
4. College of Life Sciences, Yantai University, Yantai 264005, China