1. South China Sea Institute of Oceanology, Chinese Academy of Sciences, Guangzhou 510301, China
2. College of Marine Science, University of Chinese Academy of Sciences, Beijing 100049, China
3. Department of Oceanography and Marine Geology, University of Ruhuna, Matara 81000, Sri Lanka
4. National Aquatic Resources Research and Development Agency, Colombo 01500, Sri Lanka
5. China-Sri Lanka Belt and Road Joint Laboratory on Tropical Oceanography, Guangzhou 510301, China