1. Collaborative Innovation Center of Nanfan and High-Efficiency Tropical Agriculture, Hainan University, Sanya 572025, China
2. Key Laboratory of Quality Regulation of Tropical Horticultural Crop in Hainan Province, School of Tropical Agriculture and Forestry (School of Agricultural and Rural, School of Rural Revitalization), Hainan University, Danzhou 571700, China
3. School of Breeding and Multiplication (Sanya Institute of Breeding and Multiplication), Hainan University, Sanya 572025, China
4. Scientific Observation and Experimental Station of Tuber and Root Crops in Huang-Huai-Hai Region of Agriculture Ministry, Crop Research Institute, Shandong Academy of Agricultural Sciences, Ji’nan 250100, China