1. College of Wildlife and Protected Area, Northeast Forestry University, Harbin 150040, China
2. BGI Research, Shenzhen 518083, China
3. BGI Research, Wuhan 430074, China
4. School of Life Sciences, Heilongjiang University, Harbin 150080, China
5. State Forestry and Grassland Administration Detecting Center of Wildlife, Harbin 150040, China