1. Department of Earth and Environmental Sciences, University of Exeter, Penryn Campus, Penryn TR10 9FE, Cornwall, UK
2. Laboratory of Behavior of Lower Vertebrates, A.N. Severtsov Institute of Ecology and Evolution RAS, Moscow 119071, Russia
3. Department of Ichthyology and Hydrobiology, Saint Petersburg State University, Saint Petersburg 199034, Russia
4. Estonian Marine Institute, University of Tartu, Mäealuse, 14, 12618 Tallinn, Estonia