1. School of Ecology and Nature Conservation, Beijing Forestry University, 35 East Qinghua Road Haidian District, Beijing 100083, China
2. State Environmental Protection Key Laboratory on Biodiversity and Biosafety, Research Center for Biodiversity Conservation and Biosafety, Nanjing Institute of Environmental Sciences, Ministry of Ecology and Environment, Nanjing 210042, China
3. Key Laboratory of Ecosystem Network Observation and Modeling, Institute of Geographic Sciences and Natural Resources Research, Chinese Academy of Sciences, Beijing 100101, China