1. College of Biotechnology and Pharmaceutical Engineering, Lanzhou Jiaotong University, Lanzhou 730070, China
2. Key Laboratory of Extreme Environmental Microbial Resources and Engineering, Lanzhou 730000, China
3. Key Laboratory of Desert and Desertification, Northwest Institute of Eco-Environment and Resources, Chinese Academy of Sciences, Lanzhou 730000, China
4. State Key Laboratory of Cryospheric Sciences, Northwest Institute of Eco-Environment and Resources, Chinese Academy of Sciences, Lanzhou 730000, China
5. College of Geography and Environment Science, Northwest Normal University, Lanzhou 730070, China