1. Engineering Research Center of Marine Biological Resource Comprehensive Utilization, Third Institute of Oceanography, Ministry of Natural Resource, Xiamen 361005, China
2. Xiamen Ocean Vocational College, Xiamen 361100, China
3. Fujian Provincial Key Laboratory of Island Conservation and Development, Island Research Center, MNR, Pingtan 350400, China
4. College of Biology and Environment, Zhejiang Wanli University, Ningbo 315100, China