1. Ningxia Key Laboratory of Ruminant Molecular and Cellular Breeding, College of Animal Science and Technology, Ningxia University, Yinchuan 750021, China
2. Institute of Animal Science and Veterinary Medicine, Shandong Academy of Agricultural Sciences, No. 23788, Gongyebei Road, Jinan 250100, China
3. Shandong Ox Livestock Breeding Co., Ltd., Jinan 250100, China
4. Laboratory of Animal Genetics, Breeding and Reproduction, Ministry of Agriculture of China, National Engineering Laboratory of Animal Breeding, College of Animal Science and Technology, China Agricultural University, Beijing 100193, China
5. Beijing Consortium for Innovative Bio-Breeding, Beijing 101206, China