1. Graduate School of Integrated Science for Life, Hiroshima University, 1-4-4 Kagamiyama, Higashi-hiroshima 739-8528, Japan
2. Department of Environmental Sciences, College of Agriculture and Environmental Sciences, University of South Africa, Florida, 0-41 Calabash Building, Unisa Science Campus, cnr Pioneer Avenue and Christiaan de Wet Road, Florida 1710, Gauteng, South Africa