1. College of Marine Life Sciences, Ocean University of China, 5 Yushan Road, Qingdao 266003, China
2. School of Biological Sciences, University of East Anglia, Norwich Research Park, Norwich NR4 7TJ, UK
3. Institute of Evolution & Marine Biodiversity, Ocean University of China, 5 Yushan Road, Qingdao 266003, China
4. Laboratory for Marine Ecology and Environmental Science, Laoshan Laboratory, Qingdao 266071, China