1. College of Biology and Agriculture Resources, Huanggang Normal University, Huanggang 438000, China
2. National Key Laboratory of Agricultural Microbiology, College of Life Science and Technology, Huazhong Agricultural University, Wuhan 430070, China
3. Hubei Key Laboratory of Edible Wild Plants Conservation & Utilization, Hubei Engineering Research Center of Characteristic Wild Vegetable Breeding and Comprehensive Utilization Technology, College of Life Science, Hubei Normal University, Huangshi 435002, China