1. College of Bioscience and Biotechnology, Chubu University, Kasugai 487-8501, Aichi, Japan
2. Japan Society for the Promotion of Science, Chiyoda-ku 102-0083, Tokyo, Japan
3. Yokohama Zoo Zoorasia, Yokohama 241-0001, Kanagawa, Japan
4. Sabah Wildlife Department, Wisma Muis, Kota Kinabalu 88100, Malaysia
5. Organisms and Environment Division, Cardiff School of Biosciences, Cardiff University, Cardiff CF10 3AX, UK
6. Danau Girang Field Centre, c/o Sabah Wildlife Department, Wisma Muis, Kota Kinabalu 88100, Malaysia
7. Wildlife Research Center, Kyoto University, Sakyo-ku 606-8203, Kyoto, Japan
8. Academy of Emerging Sciences, Chubu University, Kasugai 487-8501, Aichi, Japan
9. Chubu Institute for Advanced Studies, Chubu University, Kasugai 487-8501, Aichi, Japan
10. Institute for Tropical Biology and Conservation, University Malaysia Sabah, Kota Kinabalu 88100, Malaysia