1. Department of Immunology, Tufts University School of Medicine, Boston, MA 02111, USA
2. Division of Infectious Diseases and Immunology, University of Massachusetts Chan Medical School, Worcester, MA 01605, USA
3. Section of Infectious Diseases, Boston University School of Medicine, Boston, MA 02118, USA
4. Department of Preventive Medicine and Biostatistics, F. Edward Hebert School of Medicine, Uniformed Services University, Bethesda, MD 20814, USA