1. Friedrich-Loeffler-Institut, Federal Research Institute for Animal Health, Institute of Novel and Emerging Infectious Diseases, 17493 Greifswald-Insel Riems, Germany
2. Der Gruene Zoo Wuppertal, Hubertusallee 30, 42117 Wuppertal, Germany
3. Wildlife Research Institute, State Agency for Nature, Environment and Consumer Protection North Rhine-Westphalia, Puetzchens Chaussee 228, 53229 Bonn, Germany
4. Thueringer Zoopark Erfurt, Am Zoopark 1, 99087 Erfurt, Germany