1. Jilin Provincial Institute of Education, Changchun 130022, China
2. Northeast Institute of Geography and Agroecology, Chinese Academy of Sciences, Changchun 130102, China
3. Royal Botanic Gardens Victoria, South Yarra, VIC 3141, Australia
4. Jilin Provincial Key Laboratory of Tree and Grass Genetics and Breeding, College of Forestry and Grassland Science, Jilin Agricultural University, Changchun 130118, China
5. College of Environment and Resources, Dalian Minzu University, Dalian 116600, China