1. CAS Engineering Laboratory for Yellow River Delta Modern Agriculture, Institute of Geographic Sciences and Natural Resources Research, Chinese Academy of Sciences, Beijing 100101, China
2. Key Laboratory of Genetics and Germplasm Innovation of Tropical Special Forest Trees and Ornamental Plants, Ministry of Education, College of Forestry, Hainan University, Haikou 570228, China
3. College of Resources and Environment, University of Chinese Academy of Sciences, Beijing 100049, China
4. Shandong Dongying Institute of Geographic Sciences, Dongying 257000, China