1. State Key Laboratory of Desert and Oasis Ecology, Xinjiang Institute of Ecology and Geography, Chinese Academy of Sciences, Urumqi 830011, China
2. National Fukang Station of Desert Ecosystem Ecology, Field Sciences Observation and Research Station, Chinese Academy of Sciences, Fukang 831505, China
3. University of Chinese Academy of Sciences, Beijing 100049, China
4. College of Ecology, Environment and Resources, Guangdong University of Technology, Guangzhou 510006, China