1. Department of Agronomy, College of Agronomy, Hunan Agricultural University, Changsha 410128, China
2. Southern Grain and Oil Crops Collaborative Innovation Center, Changsha 410128, China
3. Shanghai Center for Plant Stress Biology, CAS Center for Excellence in Molecular Plant Sciences, Chinese Academy of Sciences, Shanghai 200032, China
4. University of Chinese Academy of Sciences, Beijing 100049, China