1. Department of ENT, Royal National ENT & Eastman Dental Hospitals, London WC1E 6DG, UK
2. Ear Institute, University College London (UCL), London WC1X 8EE, UK
3. Medical School, University College London (UCL), London WC1E 6DE, UK
4. Division of Immunity and Infection, University College London (UCL), London WC1E 6BT, UK