1. College of Horticulture and Gardening, Yangtze University, Jingzhou 434025, China
2. State Key Laboratory of Tree Genetics and Breeding, Northeast Forestry University, Harbin 150040, China
3. Department of Biology, Genetics Institute, University of Florida, Gainesville, FL 32610, USA
4. College of Life Sciences, Qingdao Agricultural University, Qingdao 266109, China
5. Department of Biology, University of Mississippi, Oxford, MS 38677, USA