1. State Key Laboratory of Black Soils Conservation and Utilization, Northeast Institute of Geography and Agroecology, Chinese Academy of Sciences, Harbin 150081, China
2. Institute of Rural Revitalization Science and Technology, Heilongjiang Academy of Agricultural Sciences, Harbin 150028, China
3. Huma Cold Temperature Plant Germplasm Resources Protection Field Scientific Observation and Research Station of Heilongjiang Province, Northeast Institute of Geography and Agroecology, Chinese Academy of Sciences, Huma 165100, China
4. Liaoning Institute of Pomology, Yingkou 115009, China
5. College of Horticultural and Plant Protection, Inner Mongolia Key Laboratory of Wild Peculiar Vegetable Germplasm Resource and Germplasm Enhancement, Inner Mongolia Agricultural University, Hohhot 010011, China