1. Research Centre for Deep Sea and Polar Fisheries, Ocean University of China, 5 Yushan Road, Qingdao 266003, China
2. Frontiers Science Center for Deep Ocean Multispheres and Earth System (FDOMES), Ocean University of China, Qingdao 266100, China
3. College of Fisheries, Ocean University of China, 5 Yushan Road, Qingdao 266003, China