1. Guangxi Laboratory on the Study of Coral Reefs in the South China Sea, Coral Reef Research Center of China, School of Marine Sciences, Guangxi University, Nanning 530004, China
2. Nanjing Hongchuang Exploration Technology Service, Nanjing 210023, China
3. Nanjing Center, China Geological Survey, Nanjing 210016, China
4. Guangxi Academy of Sciences, Nanning 530007, China
5. Institute of Beibu Gulf Marine Industry, Fangchenggang 538000, China