1. School of Ecological and Environmental Sciences, Hainan University, Haikou 570228, China
2. School of Life Sciences, Hainan University, Haikou 570228, China
3. School of Plant Protection, Hainan University, Haikou 570228, China
4. Center for Eco-Environmental Restoration Engineering of Hainan Province, Key Laboratory of Agro-Forestry Environmental Processes and Ecological Regulation of Hainan Province, Haikou 570228, China