1. MOE Joint International Research Laboratory of Animal Health and Food Safety, College of Veterinary Medicine, Nanjing Agricultural University, Nanjing 210014, China
2. Shandong Engineering Research Center of Pig and Poultry Health Breeding and Important Disease Purification, Shandong New Hope Liuhe Co., Ltd., Qingdao 266000, China
3. Beef Cattle Industry Development Center, Fangshan 033100, China
4. College of Animal Science, Xichang University, Xichang 615012, China
5. China Agriculture Research System-Yangling Comprehensive Test Station, Xianyang 712100, China