1. Acadia Centre for Estuarine Research, Acadia University, Wolfville, NS B4P 2R6, Canada
2. Department of Mathematics and Statistics, Acadia University, Wolfville, NS B4P 2R6, Canada
3. Department of Biology, Acadia University, Wolfville, NS B4P 2R6, Canada
4. Department of Fisheries and Ocean, Dartmouth, NS B2Y 4A2, Canada
5. Fundy Ocean Research Center for Energy, Halifax, NS B3J 3N5, Canada