1. Marine Science and Technology College, Zhejiang Ocean University, Zhoushan 316022, China
2. National Engineering Research Center for Marine Aquaculture, Zhejiang Ocean University, Zhoushan 316022, China
3. School of Fisheries, Zhejiang Ocean University, Zhoushan 316022, China
4. Department of Marine Biosciences, Tokyo University of Marine Science and Technology, Tokyo 108-8477, Japan