1. Department of Botany and Microbiology, Faculty of Science, King Saud University, P.O. Box 2455, Riyadh 11451, Saudi Arabia
2. Botany and Microbiology Department, Faculty of Science, Al-Azhar University, Cairo 11884, Egypt
3. Department of Life Science and Chemistry, Constructor University, 28759 Bremen, Germany
4. Institute of Environmental Biology and Biotechnology, University of Applied Sciences Bremen, Am Neustadtwall 30, 28199 Bremen, Germany