1. Shandong Province Key Laboratory of Plant Virology, Institute of Plant Protection, Shandong Academy of Agricultural Sciences, Jinan 250100, China
2. College of Plant Protection, Collaborative Innovation Center of Fruit & Vegetable Quality and Efficient Production, Shandong Agricultural University, Taian 271018, China
3. State Key Laboratory of Agrobiotechnology, Department of Plant Pathology, China Agricultural University, Beijing 100193, China
4. State Key Laboratory of Crop Biology, College of Life Sciences, Shandong Agricultural University, Taian 271018, China